{"id":38956,"date":"2025-09-03T14:08:09","date_gmt":"2025-09-03T12:08:09","guid":{"rendered":"https:\/\/srfb.be\/?p=38956"},"modified":"2025-09-03T14:17:28","modified_gmt":"2025-09-03T12:17:28","slug":"comment-adapter-les-forets-aux-changements-climatiques","status":"publish","type":"post","link":"https:\/\/srfb.be\/en\/2025\/09\/03\/comment-adapter-les-forets-aux-changements-climatiques\/","title":{"rendered":"How can we adapt forests to climate change?"},"content":{"rendered":"<p>Climate change is having an impact on many tree species and on the very functioning of forest ecosystems. Foresters are wondering how best to adapt their forests to make them more resilient. This is no easy task, as there are no miracle recipes and the factors to be taken into account come from a variety of fields: plant physiology, soil protection, silvicultural strategy, introduction of new species and provenances, etc.<\/p>\n\n\n\n<p>This article sets out some basic points relating to these matters that foresters cannot ignore.<\/p>\n\n\n\n<div class=\"wp-block-group has-grey-color-background-color has-background is-layout-constrained wp-container-core-group-is-layout-db32e940 wp-block-group-is-layout-constrained\" style=\"padding-top:var(--wp--preset--spacing--60);padding-right:var(--wp--preset--spacing--60);padding-bottom:var(--wp--preset--spacing--60);padding-left:var(--wp--preset--spacing--60)\">\n<h3 class=\"wp-block-heading\">Summary<\/h3>\n\n\n\n<p>There is no magic formula for ensuring forest resilience. To achieve this objective, forest management must take multiple factors into account.<\/p>\n\n\n\n<p>Understanding the strategies used by tree species to adapt to drought is fundamental to making the right choice of tree species for a given site. What are avoidance strategies and tolerance strategies based on delaying desiccation? What are the advantages and limitations of these strategies? Which species are concerned?<\/p>\n\n\n\n<p>This article answers these questions about plant physiology, before going on to discuss other essential issues, including soil conservation, silvicultural techniques and the introduction of species and provenances.<\/p>\n<\/div>\n\n\n\n<h2 class=\"wp-block-heading has-vert-fonce-color-color has-text-color has-link-color wp-elements-a6708d498a95ee884b033030e044aca7\">1. Plant physiology<sup>*<\/sup><\/h2>\n\n\n\n<h3 class=\"wp-block-heading has-vert-fonce-color-color has-text-color has-link-color wp-elements-b11294c239e9893d9a58872902a29963\">Preamble<\/h3>\n\n\n\n<p>Water (raw sap<sup>1<\/sup>) rises in the shaft thanks to a tension phenomenon, directly linked to the evaporative demand of the air. This demand<br>is satisfied by evapotranspiration<sup>2<\/sup> leaves. So, under the effect of heat (solar energy), the liquid water contained in the leaves passes from<br>into the ambient air in a gaseous state. This leads to a water deficit in the leaves, which puts the tree under stress and causes the sap to rise. By a \"domino effect\", the rise in sap causes tension in the roots, which compensate by absorbing water from the soil (raw sap). <strong>Evapotranspiration is therefore the driving force behind the circulation of water in the tree.<\/strong><\/p>\n\n\n\n<p>When there is a shortage of water in the soil and drought sets in, the tension becomes greater, especially when the evaporative demand from the air is high (dry air). This can lead to ruptures in the water columns of the vessels conducting ascending sap. This rupture phenomenon is known as <strong>cavitation<\/strong>. This causes air bubbles to form in the water column.<strong>gas embolism<\/strong>. This hinders or even makes it impossible for water to circulate.<\/p>\n\n\n\n<div class=\"wp-block-media-text is-stacked-on-mobile is-vertically-aligned-center\"><figure class=\"wp-block-media-text__media\"><img loading=\"lazy\" decoding=\"async\" width=\"767\" height=\"719\" src=\"https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/photosynthese.png\" alt=\"\" class=\"wp-image-38959 size-full\" srcset=\"https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/photosynthese.png 767w, https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/photosynthese-300x281.png 300w, https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/photosynthese-13x12.png 13w, https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/photosynthese-600x562.png 600w\" sizes=\"auto, (max-width: 767px) 100vw, 767px\" \/><\/figure><div class=\"wp-block-media-text__content\">\n<h4 class=\"wp-block-heading\">Photosynthesis<\/h4>\n\n\n\n<ol class=\"wp-block-list\">\n<li>Chloroplasts trap light energy<\/li>\n\n\n\n<li>Water penetrates the leaf<\/li>\n\n\n\n<li>The CO<sub>2<\/sub> penetrates the leaf<\/li>\n\n\n\n<li>The sugar leaves the leaf<\/li>\n<\/ol>\n\n\n\n<p style=\"padding-left:var(--wp--preset--spacing--50)\">A. Vessel<br>B. Stoma<\/p>\n<\/div><\/div>\n\n\n\n<div class=\"wp-block-media-text is-stacked-on-mobile is-vertically-aligned-center\" style=\"margin-bottom:var(--wp--preset--spacing--60)\"><figure class=\"wp-block-media-text__media\"><img decoding=\"async\" src=\"https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/graphique-photosynthese-769x1024.png.webp\" alt=\"\" class=\"wp-image-38961 size-full\"\/><\/figure><div class=\"wp-block-media-text__content\">\n<p>The sugars synthesised by plants are used for respiration, growth and defence (see point 4). Some of these sugars are stored in the form of starch, mainly in the crown and the main branches.<\/p>\n<\/div><\/div>\n\n\n\n<div class=\"wp-block-group has-border-color has-grey-color-border-color is-layout-constrained wp-container-core-group-is-layout-db32e940 wp-block-group-is-layout-constrained\" style=\"border-width:3px;padding-top:var(--wp--preset--spacing--60);padding-right:var(--wp--preset--spacing--60);padding-bottom:var(--wp--preset--spacing--60);padding-left:var(--wp--preset--spacing--60)\">\n<h3 class=\"wp-block-heading\">Note<\/h3>\n\n\n\n<p>Root absorption is made possible by mycorrhizae<sup>3<\/sup>In this case, it is the fungi that pump the water from the soil before returning it to the roots. In return, the tree gives the fungi some of the sugars synthesised by photosynthesis.<\/p>\n\n\n\n<p>Some of the water in the leaves is used for photosynthesis, a photochemical reaction that transforms water (raw sap) and atmospheric carbon dioxide (absorbed by the leaves) into glucose (sugar) and produces oxygen. This complex reaction takes place in the leaves and is only possible in the presence of chlorophyll. Its simplified formula is 6 CO<sub>2<\/sub> + 6 H<sub>2<\/sub>O <span class=\"wp-rich-text-font-awesome-icon wp-font-awesome-icon\"><svg aria-hidden=\"true\" focusable=\"false\" data-prefix=\"fas\" data-icon=\"arrow-right-long\" class=\"svg-inline--fa fa-arrow-right-long\" role=\"img\" xmlns=\"http:\/\/www.w3.org\/2000\/svg\" viewbox=\"0 0 512 512\"><path fill=\"currentColor\" d=\"M502.6 278.6c12.5-12.5 12.5-32.8 0-45.3l-128-128c-12.5-12.5-32.8-12.5-45.3 0s-12.5 32.8 0 45.3L402.7 224 32 224c-17.7 0-32 14.3-32 32s14.3 32 32 32l370.7 0-73.4 73.4c-12.5 12.5-12.5 32.8 0 45.3s32.8 12.5 45.3 0l128-128z\"><\/path><\/svg><\/span> C<sub>6<\/sub>H<sub>12<\/sub>O<sub>6<\/sub> + 6 O<sub>2<\/sub>.<\/p>\n\n\n\n<p>The glucose produced is used by the plant for its metabolism<sup>4<\/sup> or stored in the form of starch (reserves).<\/p>\n<\/div>\n\n\n\n<hr class=\"wp-block-separator has-text-color has-grey-color-color has-alpha-channel-opacity has-grey-color-background-color has-background\" style=\"margin-top:var(--wp--preset--spacing--60);margin-bottom:var(--wp--preset--spacing--60)\"\/>\n\n\n\n<div class=\"wp-block-group has-small-font-size is-layout-constrained wp-block-group-is-layout-constrained\" style=\"font-style:italic;font-weight:300\">\n<ol class=\"wp-block-list\">\n<li>Water and dissolved mineral salts present in the soil.<\/li>\n\n\n\n<li>It is more correct to speak of evapotranspiration rather than transpiration, a term more suited to the animal world.<\/li>\n\n\n\n<li>Symbiotic association between a fungus and the roots of a higher plant. This is known as mycorrhizal symbiosis (symbiosis is a mutually beneficial association between several organisms).<\/li>\n\n\n\n<li>Metabolism is the set of chemical reactions that take place within each cell of a living being, enabling it to sustain life, reproduce, develop and respond to environmental stimuli. <a href=\"https:\/\/fr.wikipedia.org\/wiki\/M%C3%A9tabolisme\" target=\"_blank\" rel=\"noreferrer noopener\">Source: Wikipedia<\/a><\/li>\n<\/ol>\n\n\n\n<p style=\"padding-left:var(--wp--preset--spacing--50)\">* Plant physiology is concerned with the functioning of plant organs and tissues and seeks to clarify the nature of the mechanisms by which the organs perform their functions and the plant develops in its environment. It includes the study of water relations and photosynthesis. Source : <a href=\"https:\/\/biologievegetale.be\" target=\"_blank\" rel=\"noreferrer noopener\">https:\/\/biologievegetale.be<\/a><\/p>\n<\/div>\n\n\n\n<hr class=\"wp-block-separator has-text-color has-grey-color-color has-alpha-channel-opacity has-grey-color-background-color has-background\" style=\"margin-top:var(--wp--preset--spacing--60);margin-bottom:var(--wp--preset--spacing--60)\"\/>\n\n\n\n<h3 class=\"wp-block-heading has-vert-fonce-color-color has-text-color has-link-color wp-elements-0343475d0c5c0c71f532a071b57fed44\">1.1 Strategies of tree species to cope with drought<\/h3>\n\n\n\n<p>Among forest species, there are three strategies for adapting to drought.<\/p>\n\n\n\n<h4 class=\"wp-block-heading has-rouge-color-color has-text-color has-link-color wp-elements-83daf489fd56795a6ed7b584976f1138\">1.1.1. The avoidance strategy<\/h4>\n\n\n\n<p>The avoidance strategy is based in particular on :<\/p>\n\n\n\n<ul class=\"wp-block-list\">\n<li>the <strong>phenology<sup>1<\/sup> asynchronous<\/strong> to the dry period that occurs with certain species. In the Mediterranean region, the holm oak, for example, will grow in spring and autumn. In summer, from June to September, it stops growing altogether;<\/li>\n\n\n\n<li>the <strong>early closure of stomata<sup>2<\/sup><\/strong>, In other words, closing the stomata stops evapotranspiration and blocks water inside the plant. In other words, closing the stomata stops evapotranspiration and blocks water inside the plant.<\/li>\n<\/ul>\n\n\n\n<p>This avoidance strategy is not found in temperate hardwoods, which have limited stomatal regulation. In the event of edaphic drought, the stomata will close partially, with the degree of opening varying according to the species. The American red oak, for example, is better at this game than our native oaks.<\/p>\n\n\n\n<h4 class=\"wp-block-heading has-rouge-color-color has-text-color has-link-color wp-elements-69902f180503812a1c19a9dea86311e1\">1.1.2. Tolerance strategy by delaying desiccation<\/h4>\n\n\n\n<p>This strategy is inherent to <strong>species capable of developing deep roots<\/strong>. They are therefore likely to draw water from deep down and continue photosynthesising as long as water is available, even in the event of a \u00absurface drought\u00bb and heatwave.<\/p>\n\n\n\n<p>This strategy is also based on <strong>morphological adaptation of sap-conducting vessels<\/strong> (thicker) which increases their resistance to cavitation and therefore reduces the risk of embolism.<\/p>\n\n\n\n<h4 class=\"wp-block-heading has-rouge-color-color has-text-color has-link-color wp-elements-60c58f2bfcb052227f0927312d30c975\">1.1.3. The desiccation tolerance strategy<\/h4>\n\n\n\n<p>This strategy concerns plants capable of storing water in their tissues. It does not apply to temperate regions. Some species combine several strategies, such as holm oak, which combines avoidance and tolerance by delaying desiccation. This is also the case for certain species of Cupressus and evergreen hardwoods in general. Native oaks and Atlas cedar use the strategy of tolerance by delaying desiccation (deep rooting). Pines, on the other hand, use an avoidance strategy.<\/p>\n\n\n\n<p>Whether the strategy adopted is a single one or a combination, its effectiveness will vary from one species to another. Sessile oak, for example, will be more resistant to drought than pedunculate, which is more sensitive to lack of water and will only be at its best on sites that are constantly supplied with water.<\/p>\n\n\n\n<p>It is absolutely essential to understand these two strategies for adapting to drought in order to make the right choice of species for the site. For example, Atlas cedar is highly resistant to drought, but its root system is very sensitive to soil compactness and anaerobiosis. However, it is its powerful taproot that gives it this resistance. Consequently, it is a silvicultural mistake to plant it on soils with a compacted, waterlogged or shallow horizon. In such circumstances, the Atlas Cedar's root system does not develop, which undermines its strategy for adapting to drought. Note that Atlas cedar rooting is also severely hampered by sudden textural changes in the soil.<\/p>\n\n\n\n<hr class=\"wp-block-separator has-text-color has-grey-color-color has-alpha-channel-opacity has-grey-color-background-color has-background\" style=\"margin-top:var(--wp--preset--spacing--60);margin-bottom:var(--wp--preset--spacing--60)\"\/>\n\n\n\n<div class=\"wp-block-group has-small-font-size is-layout-constrained wp-block-group-is-layout-constrained\" style=\"font-style:italic;font-weight:300\">\n<ol class=\"wp-block-list\">\n<li>Phenology is the study of the appearance of periodic events in the living world, determined by seasonal variations in climate. Source : <a href=\"https:\/\/fr.wikipedia.org\/wiki\/Ph%C3%A9nologie\" target=\"_blank\" rel=\"noreferrer noopener\">Wikipedia<\/a><\/li>\n\n\n\n<li>Two kidney-shaped cells (stomatal cells or guard cells) rich in chlorophyll, with a small opening (ostiole) between them through which the plant exchanges gases (respiration, evapotranspiration, photosynthesis). Source : <a href=\"https:\/\/www.larousse.fr\/dictionnaires\/francais\/stomate\/74771\" target=\"_blank\" rel=\"noreferrer noopener\">Larousse dictionary<\/a>. Editor's note: stomata are generally found on the underside of leaves. In some plant species, they can also be found on the upper surface of leaves and on stems.<\/li>\n<\/ol>\n<\/div>\n\n\n\n<hr class=\"wp-block-separator has-text-color has-grey-color-color has-alpha-channel-opacity has-grey-color-background-color has-background\" style=\"margin-top:var(--wp--preset--spacing--60);margin-bottom:var(--wp--preset--spacing--60)\"\/>\n\n\n\n<h3 class=\"wp-block-heading has-vert-fonce-color-color has-text-color has-link-color wp-elements-d958bbec6f66d75e168da898d9222557\">1.2. Limitations of adaptation strategies<\/h3>\n\n\n\n<p>The closure of stomata linked to <strong>avoidance strategy<\/strong> leads to a drop in photosynthetic activity. Gas exchange between the leaf and the atmosphere is blocked, the tree preserves its water but CO<sub>2<\/sub> necessary for photosynthesis stops. The tree will therefore draw on its starch reserves to ensure its metabolism. After a one-off drought, the tree will quickly return to its normal function, but there will still be some problems. <strong>prolonged and recurrent drought<\/strong> will gradually weaken the tree (it will no longer have enough starch to produce enough new leaves in the spring), which will end up by <strong>starve to death<\/strong> (not enough photosynthesis and no more reserves).<\/p>\n\n\n\n<p>In the <strong>tolerance strategy by delaying desiccation<\/strong>, As long as water is available at depth, the tree will continue its photosynthetic activity. As soon as there is a shortage of water, embolism sets in and the tree (or branch) ends up by <strong>die of thirst<\/strong>. For species that adopt this strategy, it is the <strong>intense and prolonged drought<\/strong> that are harmful to them.<\/p>\n\n\n\n<h3 class=\"wp-block-heading has-vert-fonce-color-color has-text-color has-link-color wp-elements-6f01893a3e063f15777e8a1968e18606\">1.3 Premature leaf fall<\/h3>\n\n\n\n<p>Another possible response to drought is premature leaf fall to limit water loss (as in birch, for example). This survival strategy poses a major problem in terms of the tree's reserves, as it not only reduces photosynthetic potential but also leads to a loss of nutrients. In autumn, as the leaves fall, around 50% of the nutrients they contain are recovered and stored in the twigs for budburst the following spring. When the tree has to \u00abshed\u00bb its leaves during the growing season, there is no recovery of nutrients, and the leaves fall green to the ground. This considerably weakens the tree.<\/p>\n\n\n\n<div class=\"wp-block-group has-vert-claire-color-background-color has-background is-layout-constrained wp-container-core-group-is-layout-db32e940 wp-block-group-is-layout-constrained\" style=\"padding-top:var(--wp--preset--spacing--60);padding-right:var(--wp--preset--spacing--60);padding-bottom:var(--wp--preset--spacing--60);padding-left:var(--wp--preset--spacing--60)\">\n<h3 class=\"wp-block-heading\">Vulnerability of tree species to embolism<\/h3>\n\n\n\n<p>Vulnerability to embolism can be visualised on a graph by comparing the state of the sap in the xylem ducts.<sup>1<\/sup> (water potential in MPa) and the degree of embolism (PLC %).<\/p>\n\n\n\n<div class=\"wp-block-media-text is-stacked-on-mobile is-vertically-aligned-center\"><figure class=\"wp-block-media-text__media\"><img loading=\"lazy\" decoding=\"async\" width=\"780\" height=\"685\" src=\"https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/graphique-article.png\" alt=\"\" class=\"wp-image-38965 size-full\" srcset=\"https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/graphique-article.png 780w, https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/graphique-article-300x263.png 300w, https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/graphique-article-14x12.png 14w, https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/graphique-article-600x527.png 600w\" sizes=\"auto, (max-width: 780px) 100vw, 780px\" \/><\/figure><div class=\"wp-block-media-text__content\">\n<h4 class=\"wp-block-heading\">Vulnerability curve for a European beech<\/h4>\n\n\n\n<p class=\"has-small-font-size\" style=\"font-style:italic;font-weight:300\">Source: Wortemann R., Variabilit\u00e9 de la vuln\u00e9rabilit\u00e9 \u00e0 la cavitation chez le h\u00eatre. Xyl\u00e8me symposium, INRA Nancy, 23p.<\/p>\n<\/div><\/div>\n\n\n\n<p>A value frequently used to compare species is the P50, i.e. the water potential for which 50 % of conductivity loss is measured (embolism of 50 %). The P50 should not be considered in absolute terms, as it is standardised laboratory data. Many other parameters will come into play in nature, so the P50 is indicative but will not necessarily reflect the reality in the field.<\/p>\n\n\n\n<figure class=\"wp-block-image size-large\"><img decoding=\"async\" src=\"https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/graphique-vulnerabilite-1024x806.png.webp\" alt=\"\" class=\"wp-image-38966\" style=\"object-fit:cover\"\/><figcaption class=\"wp-element-caption\"><em>Source: Adapted from Cochard H. et al., 2016. Embolism and the vulnerability of trees to drought, INRA Clermont-Ferrand.<\/em><\/figcaption><\/figure>\n\n\n\n<p>For example, the P50 of sessile oak (Quercus petraea) is slightly higher than that of spruce (Picea abies) (see graph). If we rely solely on this figure, we can conclude that spruce is more resistant to drought than sessile oak! Things are much more complex: a sessile oak installed on deep soil with a good useful reserve (see box on page 21) will have no problems with embolism, which will not necessarily be the case with spruce (traceroot) in the event of \u00absurface drought\u00bb.\u00bb<\/p>\n\n\n\n<hr class=\"wp-block-separator has-text-color has-vert-fonce-color-color has-alpha-channel-opacity has-vert-fonce-color-background-color has-background\" style=\"margin-top:var(--wp--preset--spacing--60);margin-bottom:var(--wp--preset--spacing--60)\"\/>\n\n\n\n<div class=\"wp-block-group has-small-font-size is-layout-constrained wp-block-group-is-layout-constrained\" style=\"font-style:italic;font-weight:300\">\n<ol class=\"wp-block-list\">\n<li>The part of the wood in which the raw sap circulates.<\/li>\n<\/ol>\n<\/div>\n<\/div>\n\n\n\n<h3 class=\"wp-block-heading has-vert-fonce-color-color has-text-color has-link-color wp-elements-43bb3bddf4437246e3c30f559be1360e\">1.4. Defence and water stress<\/h3>\n\n\n\n<p>The synthesis of trees' defence molecules (polyphenols, alkaloids, etc.) requires an enormous amount of energy (sugars) (see diagram in preamble). In the event of water stress, which leads to a drop in photosynthesis and therefore a reduction in reserves, the tree will give priority to reducing the most energy-intensive activity, i.e. the synthesis of defence molecules, in order to ensure its metabolic survival (respiration and growth).<\/p>\n\n\n\n<p>This principle obviously varies from species to species. Spruce is a typical example of the negative effect of water stress on the species' defences. In the event of water stress, the spruce will stop synthesising its defence molecules. The result, which is unfortunately well known to foresters, is massive attacks by the spruce bark beetle in monospecific stands.<\/p>\n\n\n\n<h3 class=\"wp-block-heading has-vert-fonce-color-color has-text-color has-link-color wp-elements-009aa3a7dda53e8df88ec621373e22db\">1.5 Indicators of tree decline<\/h3>\n\n\n\n<p>Trees that have suffered one or more droughts show the following symptoms:<\/p>\n\n\n\n<ol class=\"wp-block-list\">\n<li>branch or tree mortality, depending on the degree of cavitation within the individuals ;<\/li>\n\n\n\n<li>sparse crowns at budburst in the year following stress;<\/li>\n\n\n\n<li>a reduction in growth ;<\/li>\n\n\n\n<li>sensitivity to pathogens.<\/li>\n<\/ol>\n\n\n\n<h3 class=\"wp-block-heading has-vert-fonce-color-color has-text-color has-link-color wp-elements-91e81e7b8836717520c5e6820a2c16d9\">1.6. Growth and adaptation strategy<\/h3>\n\n\n\n<p>It is logical that droughts should have a different impact on tree growth depending on their adaptation strategy. The avoidance strategy will have a negative impact on growth (reduced photosynthesis). On the other hand, a pedunculate oak can perfectly well maintain its productivity in the middle of a drought if its roots are sufficiently deep and find available water (no drop in photosynthesis).<\/p>\n\n\n\n<p>Furthermore, installation on the best sites from a hydric point of view will not have as positive an impact in terms of growth in species that use the desiccation delay tolerance strategy as in species that rely on the avoidance strategy. The former are genetically \u00abprogrammed\u00bb to \u00abarmour\u00bb their vessels to resist cavitation, which requires a lot of energy and carbon. They will always \u00abconcentrate\u00bb on this aspect first. The latter, on the other hand, freed from water constraints, will photosynthesise in favour of their growth. In other words, the allocation of sugars produced by photosynthesis has different priorities depending on the species' adaptation strategy.<\/p>\n\n\n\n<p>Differences in growth gain depending on the site are also marked for species that use the same strategy. For example, a pubescent oak will have a narrower range of potential growth than a sessile oak, because the pubescent oak has a stronger strategy of tolerance by delaying desiccation. In other words, the pubescent oak will not grow as much (or at all) as the sessile oak on the best sites, but will be much more resistant to drought on difficult sites.<\/p>\n\n\n\n<div class=\"wp-block-columns has-vert-claire-color-background-color has-background has-link-color wp-elements-2bfe4c7f19085c056c1d0675def96e80 is-layout-flex wp-container-core-columns-is-layout-61ccdecd wp-block-columns-is-layout-flex\" style=\"padding-top:var(--wp--preset--spacing--60);padding-right:var(--wp--preset--spacing--60);padding-bottom:var(--wp--preset--spacing--60);padding-left:var(--wp--preset--spacing--60)\">\n<div class=\"wp-block-column is-layout-flow wp-block-column-is-layout-flow\" style=\"flex-basis:50%\">\n<h2 class=\"wp-block-heading\">Useful soil water reserve<\/h2>\n\n\n\n<p>The useful reserve is the quantity of water that can be used by plants within the thickness of soil that can be exploited by the roots. It is expressed in millimetres. It depends on the type of rooting of the species and the characteristics of the soil. It is directly proportional to the depth of exploitable soil. The texture of the soil (clay, silt, sand) is fundamental. Clay soils will have a higher useful reserve than silty soils, and even more so than sandy soils. The useful reserve will also be influenced by the load of coarse elements (gravel &gt; 2 mm, pebbles). The higher the load, the lower the useful reserve.<br>On average, one hectare of temperate forest evapotranspires 30 to 40 tonnes of water per day, i.e. a water depth of 3 to 4 mm.<\/p>\n<\/div>\n\n\n\n<div class=\"wp-block-column is-layout-flow wp-block-column-is-layout-flow\" style=\"flex-basis:50%\">\n<h2 class=\"wp-block-heading\">Importance of air humidity (hygrometry)<\/h2>\n\n\n\n<p>Some species are naturally adapted to atmospheric conditions with a high level of humidity. This is particularly true of certain American coastal species, where the higher the atmospheric humidity, the lower the demand for soil water. These include heterophyllous tsuga and western red cedar. Hygrometry is a factor not to be overlooked when choosing species (altitude, topography, etc.). Always consult the species sheets in the Ecological Species File (<a href=\"https:\/\/fichierecologique.be\" target=\"_blank\" rel=\"noreferrer noopener\">https:\/\/fichierecologique.be<\/a>).<\/p>\n<\/div>\n<\/div>\n\n\n\n<h3 class=\"wp-block-heading has-vert-fonce-color-color has-text-color has-link-color wp-elements-99e4dc9dca89f86fab903d244877a24a\">1.7 Rooting stresses<\/h3>\n\n\n\n<p>It is the roots located at a depth of more than one metre that ensure the absorption of water in dry periods. The graphs below show that 80 % of the roots are located before 60 cm, 12 % between 60 cm and 1 m and only 8 % beyond the metre (study carried out on ash). In spring, most of the \u00absurface\u00bb roots absorb the water needed by the tree, but as the growing season progresses (less water available at shallow depths), the deep roots take over and end up absorbing almost all the water.<\/p>\n\n\n\n<p>Consequently, rooting constraints are of major importance and undoubtedly influence the choice of species to be planted on a given site (see photos below).<\/p>\n\n\n\n<p>The main constraints on rooting are waterlogging (anaerobiosis) and soil compactness. Sensitivity to these two factors varies from species to species. Sessile oak, for example, is sensitive to prolonged waterlogging but not very sensitive to soil compactness. Atlas cedar is very sensitive to both.<\/p>\n\n\n\n<figure class=\"wp-block-image size-large\"><img loading=\"lazy\" decoding=\"async\" width=\"1024\" height=\"376\" src=\"https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/quatre-graphique-1024x376.png\" alt=\"\" class=\"wp-image-38973\" srcset=\"https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/quatre-graphique-1024x376.png 1024w, https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/quatre-graphique-300x110.png 300w, https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/quatre-graphique-18x7.png 18w, https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/quatre-graphique-600x220.png 600w\" sizes=\"auto, (max-width: 1024px) 100vw, 1024px\" \/><figcaption class=\"wp-element-caption\"><em>Changes in the quantity of water withdrawn from the soil during the summer season as a function of depth\ufffedeur (Ash). Adapted from Br\u00e9da N. et al. 2002.<\/em><\/figcaption><\/figure>\n\n\n\n<div class=\"wp-block-columns is-layout-flex wp-container-core-columns-is-layout-28f84493 wp-block-columns-is-layout-flex\">\n<div class=\"wp-block-column is-layout-flow wp-block-column-is-layout-flow\">\n<figure class=\"wp-block-image size-full\"><img loading=\"lazy\" decoding=\"async\" width=\"659\" height=\"951\" src=\"https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/gauche.png\" alt=\"\" class=\"wp-image-38974\" srcset=\"https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/gauche.png 659w, https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/gauche-208x300.png 208w, https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/gauche-8x12.png 8w, https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/gauche-600x866.png 600w\" sizes=\"auto, (max-width: 659px) 100vw, 659px\" \/><\/figure>\n<\/div>\n\n\n\n<div class=\"wp-block-column is-layout-flow wp-block-column-is-layout-flow\">\n<figure class=\"wp-block-image size-full\"><img loading=\"lazy\" decoding=\"async\" width=\"659\" height=\"951\" src=\"https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/centre.png\" alt=\"\" class=\"wp-image-38975\" srcset=\"https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/centre.png 659w, https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/centre-208x300.png 208w, https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/centre-8x12.png 8w, https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/centre-600x866.png 600w\" sizes=\"auto, (max-width: 659px) 100vw, 659px\" \/><\/figure>\n<\/div>\n\n\n\n<div class=\"wp-block-column is-layout-flow wp-block-column-is-layout-flow\">\n<figure class=\"wp-block-image size-full\"><img loading=\"lazy\" decoding=\"async\" width=\"659\" height=\"951\" src=\"https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/droite.png\" alt=\"\" class=\"wp-image-38976\" srcset=\"https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/droite.png 659w, https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/droite-208x300.png 208w, https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/droite-8x12.png 8w, https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/droite-600x866.png 600w\" sizes=\"auto, (max-width: 659px) 100vw, 659px\" \/><\/figure>\n<\/div>\n<\/div>\n\n\n\n<p>Good illustrations of the correlation between drought adaptation strategies and rooting constraints. On the left, a 12-year-old Atlas cedar (drought tolerance strategy) on a cracked slab. After a few difficult years, the cedar's roots have found their way into the cracks and are growing deep down to ensure a supply of water. In the centre, an Atlas cedar of the same age on soil with a compact slab at 40 cm. Its growth is almost non-existent, and the annual shoots are drying out. Contrary to popular belief, the cedar does not regulate its evapotranspiration very well and has very poor drought resistance. It makes up for this with its deep roots, provided they can develop. On the right, 12-year-old Arizona cypresses on the same compact slab soil. Arizona cypress uses a mixed strategy. If it cannot develop its roots (delaying strategy), it limits its evapotranspiration by closing its stomata (avoiding strategy). <em>Photos : <a href=\"https:\/\/onf.fr\/\" target=\"_blank\" rel=\"noreferrer noopener\">National Forestry Office (ONF)<\/a><\/em>.<\/p>\n\n\n\n<h2 class=\"wp-block-heading has-vert-fonce-color-color has-text-color has-link-color wp-elements-6d728f724b5cbf2b311fe386cbaf63d0\">2. Soil protection<\/h2>\n\n\n\n<h3 class=\"wp-block-heading has-vert-fonce-color-color has-text-color has-link-color wp-elements-2b4ba3d72f3daca52bfa7e16602f653a\">2.1. Nutrient export<\/h3>\n\n\n\n<p>The shorter the rotation, the greater the export of nutrients (nitrogen, phosphorus, potassium) from the forest. In fact, the higher the proportion of juvenile wood in the holdings, the greater the export of nutrients.<\/p>\n\n\n\n<p>The graph below provides essential information about the risk of soil impoverishment as a result of forestry operations. The vast majority of nutrients (50 to 80 % depending on the species) are found in branches with a diameter of less than 7 centimetres.<\/p>\n\n\n\n<p>Consequently, exporting fine branches should be avoided at all costs, especially on poor soils, where harvesting by the whole tree (including the crown) should be avoided.<\/p>\n\n\n\n<p>One of the silvicultural adaptations put forward in the face of climate change is to shorten silvicultural cycles, but beware of exports: shortening revolutions without limiting them will have disastrous effects on our forest soils, our primary production tool. The cumulative effect of shortening rotations and exploiting the total biomass leads to very high exports that the natural mineral balance of an ecosystem cannot compensate for.<\/p>\n\n\n\n<p>Such exports impoverish the soil, which becomes more acidic. This has a negative impact on :<\/p>\n\n\n\n<ul class=\"wp-block-list\">\n<li>structural stability: the soil becomes more susceptible to erosion;<\/li>\n\n\n\n<li>biological activity in the soil ;<\/li>\n\n\n\n<li>productivity.<\/li>\n<\/ul>\n\n\n\n<div class=\"wp-block-group has-border-color has-grey-color-border-color is-layout-constrained wp-container-core-group-is-layout-db32e940 wp-block-group-is-layout-constrained\" style=\"border-width:3px;padding-top:var(--wp--preset--spacing--60);padding-right:var(--wp--preset--spacing--60);padding-bottom:var(--wp--preset--spacing--60);padding-left:var(--wp--preset--spacing--60)\">\n<h3 class=\"wp-block-heading\">Note<\/h3>\n\n\n\n<ul class=\"wp-block-list\">\n<li>Windrowing slash is not a good idea, as the nutrients will be released locally rather than over the whole plot. However, for economic reasons, this practice is preferred to shredding.<\/li>\n\n\n\n<li>It is easy to understand that coppicing is the type of silvicultural treatment that has the greatest impact on exports, especially in short rotation. Old coppicing practices have ruined many forest soils.<\/li>\n\n\n\n<li>Wood left on the ground represents a significant water reserve that is often overlooked. Dead wood retains large quantities of water, some of which will be returned to the soil in the event of water stress.<\/li>\n\n\n\n<li>Putting wood-energy ash back into the forest is a false solution, because wood ash no longer contains nitrogen and carbon. These two elements are essential for tree growth and biological activity in the soil.<\/li>\n<\/ul>\n<\/div>\n\n\n\n<div class=\"wp-block-group has-vert-claire-color-background-color has-background is-layout-constrained wp-container-core-group-is-layout-a9681907 wp-block-group-is-layout-constrained\" style=\"margin-top:var(--wp--preset--spacing--50);margin-bottom:var(--wp--preset--spacing--50);padding-top:var(--wp--preset--spacing--60);padding-right:var(--wp--preset--spacing--60);padding-bottom:var(--wp--preset--spacing--60);padding-left:var(--wp--preset--spacing--60)\">\n<h3 class=\"wp-block-heading\">Hardwood or softwood<\/h3>\n\n\n\n<p>Given the climatic context, on soils with a xeric tendency<sup>1<\/sup> Hardwood forestry is being called into question to a greater extent than softwood forestry, because softwoods offer a wider range of species capable of withstanding drought, while at the same time offering high productivity.<\/p>\n\n\n\n<p>As a result, foresters are faced with a paradox: our society is more inclined to favour hardwoods than softwoods, and even tends to denigrate the latter, which are sometimes accused of all the evils. It is interesting to note that in the Mediterranean region, production forests are all made up of softwoods (Aleppo pine, maritime pine, umbrella pine, etc.).<\/p>\n\n\n\n<hr class=\"wp-block-separator has-text-color has-vert-fonce-color-color has-alpha-channel-opacity has-vert-fonce-color-background-color has-background\" style=\"margin-top:var(--wp--preset--spacing--60);margin-bottom:var(--wp--preset--spacing--60)\"\/>\n\n\n\n<div class=\"wp-block-group has-small-font-size is-layout-constrained wp-block-group-is-layout-constrained\" style=\"font-style:italic;font-weight:300\">\n<ol class=\"wp-block-list\">\n<li>Refers to an environment characterised by a very poor water balance leading to marked drought. Source: Vocabulaire forestier - \u00c9cologie, gestion et conservation des espaces bois\u00e9s, Christian Gauberville, Yves Bastien, CNPF-IDF - 2011.<\/li>\n<\/ol>\n<\/div>\n<\/div>\n\n\n\n<h3 class=\"wp-block-heading has-vert-fonce-color-color has-text-color has-link-color wp-elements-9564b3cb074e183e334f431d37a959f7\">2.2. Avoid soil compaction<\/h3>\n\n\n\n<p>Foresters should be aware that :<\/p>\n\n\n\n<ul class=\"wp-block-list\">\n<li>the porosity of the soil is of biological origin. Earthworms, in particular, aerate the soil. This aeration allows water and air to circulate in the soil;<\/li>\n\n\n\n<li>Settling is \u00abdefinitive\u00bb (see note), and there is no mechanical way of remedying it;<\/li>\n\n\n\n<li>settling reduces productivity;<\/li>\n\n\n\n<li>repeated use of heavy machinery on soil, even dry soil, leads to structural deterioration<sup>1<\/sup>;<\/li>\n\n\n\n<li>Stands on compacted soils are more rapidly weakened in the event of stress, and dieback is consequently more frequent. The loss of porosity leads to numerous problems such as waterlogging, poor rooting, a reduction in mycorrhizae, a loss of biological activity in the soil, etc.<\/li>\n<\/ul>\n\n\n\n<p>Consequently, certain recommendations are imperative in order to minimise and circumscribe the passage of heavy machinery:<\/p>\n\n\n\n<ul class=\"wp-block-list\">\n<li>Compulsory operating partitions and compliance with these in the specifications;<\/li>\n\n\n\n<li>encourage cabling of logs (wider partitions) ;<\/li>\n\n\n\n<li>Work the soil sparingly and occasionally (only when vegetation is blocked). Avoid touching the soil (e.g. by milling) to avoid destructuring it (soil life is mainly present in the first 15-20 centimetres) and mineralisation (leaching into watercourses);<\/li>\n<\/ul>\n\n\n\n<div class=\"wp-block-group has-border-color has-grey-color-border-color is-layout-constrained wp-container-core-group-is-layout-db32e940 wp-block-group-is-layout-constrained\" style=\"border-width:3px;padding-top:var(--wp--preset--spacing--60);padding-right:var(--wp--preset--spacing--60);padding-bottom:var(--wp--preset--spacing--60);padding-left:var(--wp--preset--spacing--60)\">\n<h3 class=\"wp-block-heading\">Note<\/h3>\n\n\n\n<p>Working the soil on the surface will help the plants to take root, using a reversible scarifier for example. But deep soil compaction persists, even with subsoiling. It lasts longer than the human timescale. Certain tree species can help the soil to recover by decompacting it with their roots. Adding organic matter to encourage soil biology is an essential element in the remediation of compacted soils.<\/p>\n<\/div>\n\n\n\n<hr class=\"wp-block-separator has-text-color has-grey-color-color has-alpha-channel-opacity has-grey-color-background-color has-background\" style=\"margin-top:var(--wp--preset--spacing--60);margin-bottom:var(--wp--preset--spacing--60)\"\/>\n\n\n\n<div class=\"wp-block-group has-small-font-size is-layout-constrained wp-block-group-is-layout-constrained\" style=\"font-style:italic;font-weight:300\">\n<ol class=\"wp-block-list\">\n<li>Lamand\u00e9 M. et al., 2005. Effects of forest exploitation on soil quality. ONF - Collection dossiers forestiers n\u00b015, 131p<\/li>\n<\/ol>\n<\/div>\n\n\n\n<hr class=\"wp-block-separator has-text-color has-grey-color-color has-alpha-channel-opacity has-grey-color-background-color has-background\" style=\"margin-top:var(--wp--preset--spacing--60);margin-bottom:var(--wp--preset--spacing--60)\"\/>\n\n\n\n<h2 class=\"wp-block-heading has-vert-fonce-color-color has-text-color has-link-color wp-elements-0abbe2d6f89553b83d162589e73a15f2\">3. Silvicultural strategy<\/h2>\n\n\n\n<h3 class=\"wp-block-heading has-vert-fonce-color-color has-text-color has-link-color wp-elements-21a5a4df72b0e740693d299362ec4cae\">3.1 How can forestry be adapted to climate change?<\/h3>\n\n\n\n<p>As well as introducing new species and provenances, and using the silvicultural levers discussed below, we also need to ensure that we allow the local resource to express its capacity to adapt, and not consider that the \u00abgame is up\u00bb. The intrinsic diversity of trees is very high, at least in certain species, which gives them great capacity for adaptation.<\/p>\n\n\n\n<p>To adapt forests to climate change, foresters need to act on three levels: density, structure and composition. Density refers to the intensity and frequency of thinning. The multi-storey structure refers to a layered forest with trees of multiple ages (irregular high forest). Finally, composition refers to the mixture of species (mixed forest).<\/p>\n\n\n\n<h4 class=\"wp-block-heading has-rouge-color-color has-text-color has-link-color wp-elements-f68e4b0447382b108c07555acfaf7616\">Acting on density<\/h4>\n\n\n\n<p>A more dynamic thinning regime tends to improve the water balance because, on the one hand, there is less interception of rain by the foliage, which increases the quantity of water that reaches the ground before being evaporated, and on the other hand, there is less competition for the water resource because there are fewer trees (less evapotranspiration). In other words, the denser the stand, the faster the soil water deficit threshold is reached. Thinned stands grow better during a drought<sup>1<\/sup> and better growth resilience after a dry spell.<\/p>\n\n\n\n<p>However, there are limits to the intensity of intervention, as thinning can harm residual trees. The stronger the thinning, the better the water balance in the plot, but evapotranspiration at tree level increases (development of the crown and in full light).<\/p>\n\n\n\n<p>This is an important fact to bear in mind, particularly in continuous cover mixed silviculture (CCMS). Objective trees that have been cropped will be more exposed to water stress. In CCM, even more than for other silvicultural treatments, care must be taken to ensure that the species\/station match is optimal from a hydric point of view, all the more so if we designate shade species (beech, for example) whose leaves evapotranspire abundantly. You should also be aware that the light given to the lower strata encourages their growth, which will have a significant impact on the water balance of the stand as a whole.<\/p>\n\n\n\n<p>Dynamic silviculture is fa\ufffevorable to the water supply of the stand on condition that it is not :<\/p>\n\n\n\n<ul class=\"wp-block-list\">\n<li>nor excessive ;<\/li>\n\n\n\n<li>or brutal;<\/li>\n\n\n\n<li>and early (and do not neglect depressions).<\/li>\n<\/ul>\n\n\n\n<p>The resulting reduction in the number of revolutions makes it possible to :<\/p>\n\n\n\n<ul class=\"wp-block-list\">\n<li>limit the ageing of stands, which increases their vulnerability to all types of stress ;<\/li>\n\n\n\n<li>limit exposure to risks over time ;<\/li>\n\n\n\n<li>Promote wood production for mitigation (sequestration and substitution).<\/li>\n<\/ul>\n\n\n\n<p>This is subject to vigilance regarding exports (see point 1 of the previous chapter).<\/p>\n\n\n\n<h4 class=\"wp-block-heading has-rouge-color-color has-text-color has-link-color wp-elements-b17bbb38f044b21f194f274c16266404\">Acting on the structure<\/h4>\n\n\n\n<p>On plains and soils regularly subject to drought, it has not been proven that irregular high forest is one of the forest management solutions that can limit water stress. It is not possible to say that one system is better than the other in these conditions.<\/p>\n\n\n\n<p>Natural stands that are regularly subjected to drought are regular, such as forests of Aleppo pine or ponderosa pine.<\/p>\n\n\n\n<p>Can we conclude from this that on soils with a xeric tendency, the irregular structure can be called into question? This question in no way calls into question the use of irregular high forest on less constraining and, a fortiori, optimal sites.<\/p>\n\n\n\n<h4 class=\"wp-block-heading has-rouge-color-color has-text-color has-link-color wp-elements-3147a1fcd2dfe7217fb23c1882ce6ca5\">Acting on the composition<\/h4>\n\n\n\n<p>Mixing species has positive and negative interactions.<br>Negative interactions are :<\/p>\n\n\n\n<ul class=\"wp-block-list\">\n<li>competition for resources between species in the same ecological niche<sup>1<\/sup>;<\/li>\n\n\n\n<li>antagonism by allelopathy<sup>2<\/sup>.<\/li>\n<\/ul>\n\n\n\n<p>Positive interactions are :<\/p>\n\n\n\n<ul class=\"wp-block-list\">\n<li>Complementarity: the available resources will be better shared between species with different ecological niches (tracing\/pivoting roots, shade\/light species, etc.);<\/li>\n\n\n\n<li>facilitation: the beneficial effect of one species on another species with which it interacts.<br>A well-known example is the Robinia\/Poplar association, which is beneficial to the poplar, which grows faster because of the nitrogen fixation by the Robinia. Another example is the alder\/pedunculate oak association on temporary groundwater. The alder buds earlier than the pedunculate oak and dries out the soil (evapotranspiration), which makes it easier for the oak to develop deep roots, making it more resistant to summer drought.<\/li>\n<\/ul>\n\n\n\n<p>In practice, the interactions due to diversity are positive for the level of exposure to drought (lower) for temperate deciduous forests on deep soil.<sup>3<\/sup>. The sharing of underground space by roots seems to play an important role in the effect of diversity on drought resistance.<sup>4<\/sup>.<\/p>\n\n\n\n<div class=\"wp-block-group has-border-color has-grey-color-border-color is-layout-constrained wp-container-core-group-is-layout-db32e940 wp-block-group-is-layout-constrained\" style=\"border-width:3px;padding-top:var(--wp--preset--spacing--60);padding-right:var(--wp--preset--spacing--60);padding-bottom:var(--wp--preset--spacing--60);padding-left:var(--wp--preset--spacing--60)\">\n<h3 class=\"wp-block-heading\">Note<\/h3>\n\n\n\n<p>In Mediterranean conditions, the level of exposure to drought is controlled not by diversity but by stem density.<\/p>\n<\/div>\n\n\n\n<h5 class=\"wp-block-heading has-vert-fonce-color-color has-text-color has-link-color wp-elements-6861f2023af6c7f29390774bb5ecf250\">Composition and growth<\/h5>\n\n\n\n<p>In terms of growth, mixed stands are generally more productive (particularly in terms of improving hay). The figures below give us the following information (left figure = overall effect of mixing, right figure = effect of mixing by species):<\/p>\n\n\n\n<ul class=\"wp-block-list\">\n<li>in the beech\/oak mixture, the effect is slightly positive for the beech, but the oak loses out. The oak will have more vitality in a pure oak forest than in a beech\/oak forest.<\/li>\n\n\n\n<li>the beech\/spruce mixture is very favourable to the growth of the beech, but not to that of the spruce;<\/li>\n\n\n\n<li>the beech\/fir mixture is particularly favourable to the growth of the beech;<\/li>\n\n\n\n<li>the spruce\/hemlock mix is favourable to the spruce;<\/li>\n\n\n\n<li>for the oak\/pine mixture, the mixture effect is positive for oak but not for pine;<\/li>\n<\/ul>\n\n\n\n<div class=\"wp-block-media-text is-stacked-on-mobile is-vertically-aligned-center\" style=\"grid-template-columns:75% auto\"><figure class=\"wp-block-media-text__media\"><img decoding=\"async\" src=\"https:\/\/srfb.be\/wp-content\/uploads\/2025\/09\/graphiqe-peuplement-1024x543.png.webp\" alt=\"\" class=\"wp-image-38982 size-full\"\/><\/figure><div class=\"wp-block-media-text__content\">\n<p class=\"has-small-font-size\" style=\"font-style:italic;font-weight:300\">Adapted from To\u00efgo M., et al. 2015. Overyielding in mixed forests decreases with site productivity. Journal of Ecology (103) pp. 502-512<\/p>\n<\/div><\/div>\n\n\n\n<hr class=\"wp-block-separator has-text-color has-grey-color-color has-alpha-channel-opacity has-grey-color-background-color has-background\" style=\"margin-top:var(--wp--preset--spacing--60);margin-bottom:var(--wp--preset--spacing--60)\"\/>\n\n\n\n<div class=\"wp-block-group has-small-font-size is-layout-constrained wp-block-group-is-layout-constrained\" style=\"font-style:italic;font-weight:300\">\n<ol class=\"wp-block-list\">\n<li>All the parameters that characterise the ecological requirements or lifestyle of a species. Source: Vocabulaire forestier - \u00c9cologie, gestion et conservation des espaces bois\u00e9s, Christian Gauberville, Yves Bastien, CNPF-IDF - 2011.<\/li>\n\n\n\n<li>Allelopathy is a biological phenomenon in which an organism produces one or more biochemical substances that influence the germination, growth, survival and reproduction of other organisms. Source : <a href=\"https:\/\/fr.wikipedia.org\/wiki\/All%C3%A9lopathie\" target=\"_blank\" rel=\"noreferrer noopener\">Wikipedia<\/a>.<\/li>\n\n\n\n<li>Grossiord C., et al., 2015. Temperate forests facing the consequences of climate change. RFF 2, pp. 99-110.<\/li>\n\n\n\n<li>Grossiord C., et al., 2015. Temperate forests facing the consequences of climate change. RFF 2, pp. 99-110.<\/li>\n<\/ol>\n<\/div>\n\n\n\n<hr class=\"wp-block-separator has-text-color has-grey-color-color has-alpha-channel-opacity has-grey-color-background-color has-background\" style=\"margin-top:var(--wp--preset--spacing--60);margin-bottom:var(--wp--preset--spacing--60)\"\/>\n\n\n\n<h2 class=\"wp-block-heading has-vert-fonce-color-color has-text-color has-link-color wp-elements-9fd05a7441cc9197df41d536054ba48d\">4. Introduction of new species<\/h2>\n\n\n\n<p>It's worth remembering that the European continent lost a huge number of species during the ice ages, which is not the case for North America.<sup>1<\/sup>. Thus, at comparable latitudes, it is not uncommon to find a ten-to-one ratio in terms of species in favour of the American continent.<\/p>\n\n\n\n<p>The aim of introducing new species and provenances is to maintain forestry, particularly production forestry, with species that are better adapted to drier and warmer conditions.<sup>2<\/sup>. It is based on assisted migration (see below) and the introduction of non-native species (see box). These introductions require an in-depth knowledge of the autecology of the species.<sup>3<\/sup> essences.<\/p>\n\n\n\n<p>The Trees For Future project<sup>4<\/sup>, conducted by the Soci\u00e9t\u00e9 Royale Foresti\u00e8re de Belgique, is testing different tree species and provenances in forests within a network of experimental plots spread across the country (see article on page 8).<\/p>\n\n\n\n<h4 class=\"wp-block-heading has-rouge-color-color has-text-color has-link-color wp-elements-d647f7edb48376f203614ac9b98c967e\">Assisted provenance migration<\/h4>\n\n\n\n<p>This involves introducing trees of a species that is already present in the area but from the south of its range. For example, if a common beech is planted in Belgium from seed harvested in Italy or the south of France, this is known as assisted migration of provenances. The aim here is to introduce drought resistance genes present in these southern populations and to allow them to be propagated within the local populations. The aim is therefore to enrich the genetic diversity of a local species to enable it to adapt to the new climate. The species concerned by this type of migration in the Trees for Future project are the sessile oak, the beech, the small-leaved lime and, to a lesser extent, the downy oak because, although indigenous, it is very rare here today.<\/p>\n\n\n\n<h4 class=\"wp-block-heading has-rouge-color-color has-text-color has-link-color wp-elements-2d3ba625e5ecd2e3b09c828e51bf4e74\">Assisted species migration<\/h4>\n\n\n\n<p>In this case, we are introducing a species that is not yet present in Belgium but which lives further south on the continent in climatic conditions similar to those expected in Belgium over the coming decades. For example, when Mediterranean oaks (Hungarian oak, hairy oak), Mediterranean firs (Cephalonian fir, Bornm\u00fcller fir, etc.) or Mediterranean pines (maritime pine, Macedonian pine, Bosnian pine, etc.) are planted in Belgium, this is referred to as assisted species migration.<\/p>\n\n\n\n<div class=\"wp-block-group has-orange-color-background-color has-background is-layout-constrained wp-container-core-group-is-layout-db32e940 wp-block-group-is-layout-constrained\" style=\"padding-top:var(--wp--preset--spacing--60);padding-right:var(--wp--preset--spacing--60);padding-bottom:var(--wp--preset--spacing--60);padding-left:var(--wp--preset--spacing--60)\">\n<h3 class=\"wp-block-heading\">Towards a continental definition of indig\u00e9nat?<\/h3>\n\n\n\n<p class=\"has-small-font-size\">Box extracted from the article \u00abIntroducing new species and provenances in response to climate change: daring or reckless? Nicolas Dassonville and Pascaline Leruth, Silva Belgica 1\/2023.<\/p>\n\n\n\n<p>Over the course of their long existence in a given area, native tree species have developed a multitude of interactions with the local flora and fauna. It is all these interactions that give the forest ecosystem its complexity and give sub-natural old-growth forests their biological value.<sup>1<\/sup>. A species is said to be indigenous if it occurs naturally (without any recent or past intervention) and if it has been cultivated for a long time.<sup>2<\/sup>, This is the term used to describe the species (direct or indirect human activity) present in a given area. The notion of indigenousness therefore depends on the geographical scale considered. Should a species naturally present in Rochefort be considered indigenous to Famenne, Wallonia, Belgium or Western Europe? The decision is relatively arbitrary, but it can potentially have a major influence on forestry policy, particularly in terms of legislation.<\/p>\n\n\n\n<p>For example, a species considered indigenous to Flanders (e.g. Scots pine<sup>3<\/sup>) and which can therefore be planted throughout this territory, including in Natura2000 and in sub-natural old-growth forests, may be considered non-native in Wallonia, or vice versa. In the case of Scots pine, for example, a national definition of indigenousness would certainly change the situation.<\/p>\n\n\n\n<p>Given that administrative boundaries, whether national or regional, cannot be transposed to biological species, including trees, and given that climate change is progressively modifying the climatic envelopes of tree species, would it not be more appropriate to consider indigenousness at a continental level? Over the course of glacial and interglacial periods, it is likely that tree species have come into contact with each other. We can therefore consider that species from southern Europe have a history of interaction with local flora and fauna.<\/p>\n\n\n\n<p>When it comes to policies on introducing new species, shouldn't a species such as Hungarian oak be considered as indigenous to Europe and therefore not be subject to the same restrictions as a species of Asian origin (e.g. Manchurian ash) or North American origin (e.g. sequoia sempervirens) which has no history of interaction with local flora and fauna and which, a priori, has a lower biological potential and a higher invasive risk?<\/p>\n\n\n\n<hr class=\"wp-block-separator has-text-color has-vert-fonce-color-color has-alpha-channel-opacity has-vert-fonce-color-background-color has-background\" style=\"margin-top:var(--wp--preset--spacing--60);margin-bottom:var(--wp--preset--spacing--60)\"\/>\n\n\n\n<div class=\"wp-block-group has-small-font-size is-layout-constrained wp-block-group-is-layout-constrained\" style=\"font-style:italic;font-weight:300\">\n<ol class=\"wp-block-list\">\n<li>Sub-natural: describes vegetation that has a certain natural character, but masks, often incompletely, traces of past human activity (agro-pastoral, materials extraction, etc.). Source: Vocabulaire forestier. \u00c9cologie, gestion et conservation des espaces verts, Christian Gauberville, Yves Bastien - CNPF-IDF - 2011.<\/li>\n\n\n\n<li>No precise definition of time.<\/li>\n\n\n\n<li>Whether Scots pine is indigenous to Belgium is still a matter of debate. It is thought to have been sown for the first time in Belgium in 1675, in the Campine region. Source: Ecological file of tree species.<\/li>\n<\/ol>\n<\/div>\n<\/div>\n\n\n\n<div class=\"wp-block-media-text is-stacked-on-mobile is-vertically-aligned-center has-vert-claire-color-background-color has-background has-link-color wp-elements-e5f7821e75b34913e9d391087a1f8c0c\" style=\"margin-top:var(--wp--preset--spacing--60);margin-bottom:var(--wp--preset--spacing--60);padding-top:var(--wp--preset--spacing--60);padding-right:var(--wp--preset--spacing--60);padding-bottom:var(--wp--preset--spacing--60);padding-left:var(--wp--preset--spacing--60);grid-template-columns:25% auto\"><figure class=\"wp-block-media-text__media\"><img loading=\"lazy\" decoding=\"async\" width=\"1024\" height=\"802\" src=\"https:\/\/srfb.be\/wp-content\/uploads\/2024\/03\/trees-for-future-logo-couleur-edited-1024x802.png\" alt=\"Trees for Future - Logo\" class=\"wp-image-35958 size-full\" srcset=\"https:\/\/srfb.be\/wp-content\/uploads\/2024\/03\/trees-for-future-logo-couleur-edited-1024x802.png 1024w, https:\/\/srfb.be\/wp-content\/uploads\/2024\/03\/trees-for-future-logo-couleur-edited-300x235.png 300w, https:\/\/srfb.be\/wp-content\/uploads\/2024\/03\/trees-for-future-logo-couleur-edited-15x12.png 15w, https:\/\/srfb.be\/wp-content\/uploads\/2024\/03\/trees-for-future-logo-couleur-edited-600x470.png 600w, https:\/\/srfb.be\/wp-content\/uploads\/2024\/03\/trees-for-future-logo-couleur-edited.png 1537w\" sizes=\"auto, (max-width: 1024px) 100vw, 1024px\" \/><\/figure><div class=\"wp-block-media-text__content\">\n<p>On the <a href=\"https:\/\/climessences.fr\/\" target=\"_blank\" rel=\"noreferrer noopener\">https:\/\/climessences.fr\/<\/a> detailed self-ecology sheets for 120 species.<\/p>\n\n\n\n<p>The aim of these fact sheets is to provide forestry professionals with available information on native and exotic species of potential interest to French forestry.<\/p>\n\n\n\n<div class=\"wp-block-buttons is-layout-flex wp-block-buttons-is-layout-flex\">\n<div class=\"wp-block-button\"><a class=\"wp-block-button__link has-black-color has-white-background-color has-text-color has-background has-link-color wp-element-button\" href=\"https:\/\/www.treesforfuture.be\/en\/\" target=\"_blank\" rel=\"noreferrer noopener\">Visit our Trees for Future website<\/a><\/div>\n<\/div>\n<\/div><\/div>\n\n\n\n<hr class=\"wp-block-separator has-text-color has-grey-color-color has-alpha-channel-opacity has-grey-color-background-color has-background\" style=\"margin-top:var(--wp--preset--spacing--60);margin-bottom:var(--wp--preset--spacing--60)\"\/>\n\n\n\n<div class=\"wp-block-group has-small-font-size is-layout-constrained wp-block-group-is-layout-constrained\" style=\"font-style:italic;font-weight:300\">\n<p>Article from <a href=\"https:\/\/srfb.be\/en\/informations-on-forests\/silva-belgica\/\">Silva Belgica<\/a> 4\/2023<\/p>\n\n\n\n<p>Written by David Dancart<sup>1<\/sup>, based on a presentation by Jean-Claude Tissaux<sup>2<\/sup> given on the occasion of the day entitled \u00abStrategies for adapting forests to climate change\u00bb organised by the Royal Forestry Society in 2022.<\/p>\n\n\n\n<ol class=\"wp-block-list\">\n<li>Head of Silva Belgica, Royal Forestry Society of Belgium<\/li>\n\n\n\n<li>Jean-Claude Tissaux is in charge of forest reconstitution and adaptation to climate change at the Office national des for\u00eats (France). He is involved in numerous field research projects on the subject.<\/li>\n<\/ol>\n<\/div>","protected":false},"excerpt":{"rendered":"<p>Les changements climatiques impactent de nombreuses essences foresti\u00e8res et le fonctionnement m\u00eame des \u00e9cosyst\u00e8mes forestiers. Les forestiers s\u2019interrogent sur la meilleure mani\u00e8re d\u2019adapter leur for\u00eat afin de les rendre plus r\u00e9silientes. La chose n\u2019est pas ais\u00e9e car il n\u2019existe pas de recettes miracles et les facteurs \u00e0 prendre en compte \u00e9manent de domaines divers :&#8230; <\/p>\n<div class=\"clear\"><\/div>\n<p><a href=\"https:\/\/srfb.be\/en\/2025\/09\/03\/comment-adapter-les-forets-aux-changements-climatiques\/\" class=\"excerpt-read-more\">Read More<\/a><\/p>","protected":false},"author":18,"featured_media":38988,"comment_status":"closed","ping_status":"closed","sticky":false,"template":"","format":"standard","meta":{"iawp_total_views":301,"footnotes":""},"categories":[7],"tags":[],"class_list":["post-38956","post","type-post","status-publish","format-standard","has-post-thumbnail","hentry","category-actualite"],"yoast_head":"<!-- This site is optimized with the Yoast SEO plugin v27.3 - https:\/\/yoast.com\/product\/yoast-seo-wordpress\/ -->\n<title>Comment adapter les for\u00eats aux changements climatiques? - Soci\u00e9t\u00e9 Royale Foresti\u00e8re de Belgique<\/title>\n<meta name=\"description\" content=\"La r\u00e9silience foresti\u00e8re passe par une gestion int\u00e9gr\u00e9e\u00a0: choix d\u2019essences selon leur strat\u00e9gie face \u00e0 la s\u00e9cheresse, pr\u00e9servation des sols et techniques sylvicoles r\u00e9fl\u00e9chies.\" \/>\n<meta name=\"robots\" content=\"index, follow, max-snippet:-1, max-image-preview:large, max-video-preview:-1\" \/>\n<link rel=\"canonical\" href=\"https:\/\/srfb.be\/en\/2025\/09\/03\/comment-adapter-les-forets-aux-changements-climatiques\/\" \/>\n<meta property=\"og:locale\" content=\"en_GB\" \/>\n<meta property=\"og:type\" content=\"article\" \/>\n<meta property=\"og:title\" content=\"Comment adapter les for\u00eats aux changements climatiques? 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